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Wine directory structure used in Fedora Core Linux:
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===================================================
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\${HOME}/.wine/drive_c is the root directory (aka C: drive) wine looks for by
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default. It is created with (empty) C:\windows and C:\windows\system
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directories, needed to operate Wine without an existing Windows
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installation, if it doesn't already exists.
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If you want to use Wine with an existing Windows installation that is
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mounted, for example, in /mnt/windows-c, change the symlink
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\${HOME}/.wine/dosdevices/c: to point to /mnt/windows-c instead of the
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defaults set by installation.
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If you do this, you can safely remove \${HOME}/.wine/drive_c.
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(Alternatively, just mount your Windows partition to \${HOME}/.wine/drive_c.)